Tridacnid
clams are often beautiful animals, the shells
may have a pleasing architecture, and their
exposed tissues often have pleasing patterns
and bright colors. Given the appropriate conditions,
they can be quite hardy. However, clamrades,
unless they are well nourished with supplemental
feedings they are unlikely to thrive in a
captive system.
The
Tridacnids are one of the relatively few types
of clams that may be successfully kept by
aquarists. This group, which consists of the
species Hippopus hippopus and several
species in the genus Tridacna, are
maintained with moderate success by many aquarists.
These species all contain zooxanthellae in
their tissues and for good health need a lot
of light. In the real world they are seldom
found deeper than about 30 feet and the majority
of individuals are much shallower, even living
on the reef flats in only a few inches of
water. Given that they are living in the equatorial
tropics, they receive and need a VERY significant
amount of sunlight.
The
zooxanthellae are maintained in the clams'
tissues and, when sufficiently illuminated
and otherwise cared for by the clam, use the
light impinging on them and the dissolved
carbon dioxide in the clam's tissues to produce
sugars and other photosynthetic byproducts.
Aeons of natural selection have fined tuned
this association significantly. The clams
actually have clear lenses in the tissues
of their mantles to focus sunlight onto clusters
of zooxanthellae maintained deep in the clam's
tissues.
The
brightly-colored, intricately-patterned, exposed
tissue, visible between the clam's shells,
is the mantle. The clam's mantle has many
functions; it secretes the shell, has sensory
structures to allow the clam to sense the
world around it, and it creates an internal
hydrodynamic space for the water flow around
and through the filter-feeding organs, the
gills. The colored patterns seen on the shell
may have some function, perhaps as sunscreen,
but if so, that function remains obscure.
Zooxanthellae are greenish or greenish-brown,
and don't contribute much to the color patterns
seen in the clams.
While
the sugars that are by-products of their zooxanthellae
provide them with some energy, all the tridacnids
are suspension-feeding animals and get the
raw materials they need for new tissues from
the phytoplankton they eat. In fact, until
they get about four inches long, there simply
is not enough volume of the zooxanthellate
tissues for them get all of their energy requirements
from the photosynthetic sugars, and need to
get a lot of their energy, as well as raw
materials, from feeding. When they are grown
in aquaculture situations, the sea water containing
the clams is often enriched with both nitrate
and phosphate based fertilizer. As a result
of this good water and lots of sunlight, the
clams grow very rapidly. In the average reef
aquarium, phosphates and nitrates are minimized
and the light really isn't that intense. In
nature these clams seem to feed primarily
on unicellular green algae and bacteria; consequently,
in aquaria the clams benefit from frequent
additions of phytoplankton. Most tanks with
a good deep sand bed, also have a lot of bacteria
and natural microplankton in the water as
well. These will assist in clam growth. Without
supplemental feeding, Tridacnid clams typically
appear to live well for three to six months
after which they rather suddenly die. It appears
that they utilize all of their accumulate
energy reserves and in effect, live on borrowed
time. When the reserves run out, they don't
get enough nutrients from the system that
they are in and perish.
Photos courtesy of Reef Central members.
Text by Ronald L. Shimek, Ph.D.
Many thanks to John Love (Rock Anemone) for
his assistance with this project.